The remains of Ruminantia are very rare at Montredon. No specific determination was possible. We have only one Micromeryx, one small cervid, one tragocere and one (?) gazella. The faunal spectrum is in good correlation with the general retreat of the cervids in the European upper Miocene, but in contrast it is not convenient for the bovids, which are in expansion at the same time. For them, we have to invoke the local ecological conditions.
  

Carolocoutoia ferigoloi gen. et sp. nov. is the largest of protodidelphid marsupials, known from Middle Paleocene levels at Itaboraí Formation, southeastern Brazil. It differs from other members of this family in having molars with low cusps which are basally inflated, rounded crests without cutting edges, and a thick enamel layer which is wrinkled, specially at the labial half. A comparative analysis among representatives of this family led us to recognize only three genera undoubtely assignable to it:  Protodidelphis PAULA COUTO, 1952, Robertbutleria MARSHALL, 1987, and Carolocoutoia gen. nov. Protodidelphids lack the basic derived features diagnostic of Polydolopimorphian marsupials, while most of its derived features agree with its belonging to the Didelphimorphia. Protodidelphids comprise a specialized clade of opossum-like marsupials adapted to frugivorous or frugivore-omnivorous feeding habits. They differ from other didelphimorphians in having very large, spire-like entoconids, reduced and antero-posteriorly compressed paraconids, absence of stylar cusp C and of para- and metaconules, large stylar cusps B and D which are proximate to each other, short postmetacristae, eccentric protocones, and molars that increase rapidly in size from M/ml to M/m3.
  

The Paleogene faunal assemblage from Antofagasta de la Sierra (Catamarca, Argentina), is here presented, both in its geological and systematic aspects. The fossil bearing levels are referred to the Geste Formation (Pastos Grandes "Group"). The described specimens belong to the Classes Reptilia (Orders Crocodylia, Serpentes and Chelonii) and Mammalia (three taxa from the Superorder Marsupialia, representatives of the Orders Edentata, Condylarthra, Pyrotheria and Astrapotheria, and six families of the Order Notoungulata). This fauna is referred to the Mustersan Age, which in Patagonia represents the Middle Eocene. Such chronologic assignment is based on the presence of characteristic taxa, their evolutionary stage and on stratigraphic evidence. Finally, a brief comparison with other faunal assemblages from the Early Tertiary of Argentina and Chile, is presented. 

The artiodactyls (Mammalia) from the latest Ypresian locality of Prémontré from the Paris Basin (niveau repère MP 10 in the lower Eocene of the Paris Basin) are described in this paper. Three species have been identified: 1) Diacodexis cf. varleti SUDRE et al., 1983; 2) a new species of Eurodexis ERFURT & SUDRE (E. russelli nov. sp.) defined after the revision of the species Messelobunodon? ceciliensis from the Lutetian beds of Geiseltal (Germany); and 3) Eurodexeinae indet., a probable ancestor of another form from the Geiseltal which was previously recorded as Homacodon? sp. (Erfurt 1993) and now named Parahexacodus germanicus. The two later forms are referred to the new subfamily Eurodexeinae (Erfurt & Sudre 1996). The analysis of these forms as weIl as comparative studies have led us to reconsider our previous conclusions regarding the content of the species Protodichobune oweni LEMOINE 1878 and some aspects of Ypresian diacodexid evolution. One can postulate that the divergence of E. russelli nov. sp. occurred during the first radiation of these primitive artiodactyls. Some other stem form with bunodont teeth such as Protodichobune and Aumelasia have also differentiated from Diacodexis. Like Eurodexis, these two genera persist during the middle Eocene. The absence of Protodichobune and Aumelasia at Prémontré is probably due to particular ecological conditions. 

A new genus, Eppsillycteris, is erected for Adapisorex? allglicus COOPER, 1932, from the earliest Eocene Blackheath Beds of Abbey Wood, London, England. Various derived character states indicate that it belongs to the order Chiroptera (bats) rather than to the extinct "insectivore" family Adapisoricidae. Other derived character states are shared with fossil and modern members of the family Emballonuridae. Placement of the new genus in this family extends the record of the Emballonuridae back in time by about 10 million years. It is the earliest record of a modern bat family and one of the earliest bats. This implies that the differentiation of at least some modern bat families took place in the Palaeocene, where no authenticated records of bats yet exist. The primitive characters of the earliest bats make the family Nyctitheriidae an unlikely stem group for the order Chiroptera. A tentative plausible alternative exists in some unnamed upper molars from the Palaeocene of Walbeck, Germany. Wyollycteris chalix, described as a bat from the Late Palaeocene of Wyoming, U,S.A., fits better in the family Nyctitheriidae. 

Revision of the Tayassuidae from the old collections of the Quercy leads to interesting facts about taxonomic and phylogenetic problems. It could not have been well done without constant comparison with contemporaneous Tayassuidae from stratified series, as well as with European Miocene Tayassuidae and Hyotheriinae. The result is that only three species are present in the Phosphorites of Quercy: Doliochoerus quercyi FiLHOL (represented by two sub-species: the older, D. quercyi quercyi and the more recent, D. quercyi aquensis), Palaeochoerus gergorianus CROIZET et BLAINVILLE and Palaeochoerus purillus nov. sp. The two former species disappeared with Stampian time, while the third is the direct ancestor of the Aquitanian Palaeochoerus typus POMEL, which became extinct before the beginning of Burdigalian.

The comparison with Hyotheriinae brought to light new criteria for determinations, based on the study of lower premolars. These observations are condensed in a table of keys for determinations.

As a result of these studies, it came to light that the Suoidea of the Oligo-Miocene of Europe have never given rise to, except in rare cases, very long lineages. The different forms of pigs are replaced constantly in Europe by the simple game of extinctions and migrations. The reason of that discontinuity is considered here. 

Paleosols and the taphonomy of vertebrate fossils in the Eocene Kuldana Formation of northern Pakistan provide important information on the preservation and time-averaging of fossil assemblages. Morphologic, mineralogic, and chemical data as well as comparisons with Quaternary soils suggest that Kuldana paleosols formed under generally dry and oxidizing conditions over time intervals of less than 100 000 years and perhaps as short as 1000 years. The distribution of carbonate in Kuldana paleosols further indicates that the upper half of the profiles were acidic whereas the lower halves were alkaline. Vertebrate fossils are rare in Kuldana paleosols and occur primarily in well-cemented sandstones and conglomerates with abundant micritic and iron-stained nodules that were reworked from floodplain soils. The scarcity of vertebrate remains in Kuldana paleosols probably reflects a combination of acidic, dry, and oxidizing conditions in the upper half of the profiles and rapid floodplain sedimentation. Comparisons between the taphonomic characteristics of Kuldana channel fossil assemblages and bone accumulations in modem rivers provide a basis for estimating the length of time represented by Kuldana fossils from several important localities. Vertebrate fossil assemblages from Barbora Banda are characterized by a low-diversity paleofauna, partially articulated skeletons, and bones that are sorted by size and shape. Comparison with bone accumulations in modern rivers suggests that the fossils from Barbora Banda accumulated in 1 to 10 years. Vertebrate fossils from the Lower Kuldana in the Kala Chitta Hills region, typified by locality H-GSP 62, are characterized by a high-diversity paleofauna and generally random and unsorted fossil bone distributions, which suggest that the fossils from these localities represent longer time intervals than the Barbora Banda fossils. Based on the time estimates for Kuldana paleosol development, fossil assemblages in Kuldana channel deposits in the Kala Chitta Hills region probably represent time intervals of about 1000 years. 

Abstract not available 

The present work is devoted to the study of the Cretaceous and Tertiary fishes (teeth of Selachians, Actinopterygians and Dipnoans) collected during a recent expedition in Niger. The Maestrichtian localities have yielded a new genus and a new subspecies of Selachian: Igdabatis sigmodon nov. gen., nov. sp. and Lamna biauriculata nigeriana nov. subsp. The locality of Sessao, which has been attributed to the Thanetian by means of the study of the fish, has furnished by screen-washing an interesting fauna wherein six new species are described: Raja Iouisi, Dasyatis sessaoensis, D. sudrei, D. russelli, Hypolophites thaleri and Ceratodus casieri. Comparison of these faunas with contemporary faunas of Africa has brought out a certain endemism in the Iullemmeden Basin during the late Cretaceous and the early Tertiary. 

Pour la première fois un spécimen de lagomorphe a été récolté en Quercy. L'intérêt de ce fossile tient surtout à son âge géologique inattendu, qui recule considérablement la date de première apparition en Europe de cet ordre de mammifère, Ceci nous paraît justifier une nouvelle réflexion sur la « grande coupure» oligocène, 

The entelodonts of the Quercy phosphorites are essentially known from isolated teeth. The skull remains attributed to Entelodon magnum by Kowalewsky in 1876 and the type left mandible of Enteladon depereti RÉPELIN, 1918 do not belong to this family. From a morphologic point of view, the teeth from Quercy are quite comparable to those from the type localities of E. magnum and E. deguilhemi. In the absence of distinctive anatomical criteria between E. magnum (small species from the level of Ronzon) and E. deguilhemi (large species from the level of Villebramar) they are referred, according to their size, to one or the other of these two species. However, some of them are intermediate in size. Their existence speaks in favor of a close relationship between E. magnum and E. deguilhemi and confers a real stratigraphic interest to these entelodonts of Western Europe. Finally, some anatomical characters (notch on the mesial border of P4, presence of a paraconid joined to the metaconid, trigonid higher than talonid) considered until now as peculiar to the North American Archaeotherium and not yet cited in its European contemporary Entelodon have been brought to light on the teeth from Quercy.

  

The Causses" is a near 3400 km2 large plateau located in the south of France. Here the first dinosaur footprints where found in 1935. After this, this area has yielded an ever-increasing number of ichnites now in excess of 500 specimens. These latter, 15 to 50 cm long, tridactyl or tetradactyl footprints of generally biped animals, were discovered at the surface of Hettangian to lower Sinemurian dolomite layers within 4 distinct stratigraphic units. The 35 sites bearing ichnites are located on the plateau margin. For the first time, morphologic characters studied through descriptive statistic methods with the usual parameters and classical Student and Snédecor tests, allowed us, to divide the whole set of biped traces into 6 ichnospecies. Their definitions are further constrained by multivariate statistical results using Principal Component Analysis (PCA), Factor Analysis of correspondances (FAC) and Discriminant Analysis (DA). All have confirmed the morphologic observations. So that now, the following taxa are identified : Grallator variabilis, G. lescurei, G. sauclierensis, G. minusculus, Eubrontes giganteus, Dilophosauripus williamsi, cf. Moraesichnium, Orníthopus fabrei nov ichnosp. The more immediately visible differences relate to the interdigital II-IV divarication and the digit length ratio. To this panel, we must add Batrachopus deweyi and shapes suggesting Trisauropodichnus and/or Anomoepus. Among all ichnite associations described in the lower Liasic, the New England assemblage presents the most affinities with ours. It shows the ichnotaxa Grallator, Dilophosauripus, Eubrontes, Batrachopus without forgetting Ornithopus fabrei nov. ichnosp. which is close to Ornithopus gallinaceus from the Massachusetts and Connecticut basins. On comparing the present early Jurassic ichnofauna of the Causses with the ones of the Middle and Upper Triassic formations of the eastem border of the Massif Central (France), it appears that tridactyl footprints become more and more numerous and large from Triassic to Early Jurassic. In the Causses, these latest are prevalent but in Quercy (France), Poland, Italy, USA, they are also associated with Omithopoda, Thyreophora and Sauropoda ichnites. Footprint areas considered here were generaly under an arid climate. Animals that passed by were heavy and bulky possible Megalosaur trackmakers, and lighter and slender Coelophysids or Ceratosaurs. For all, these areas were pathways as the orientations of the trackways seem point out. The directions followed by these reptiles were without any important variation during the Hettango-Sinemurian stages. These areas were also used from time to time by Crocodilomorpha and may be tetradactyl (I-IV) bipedal avian Theropods. However, the number of such trackways in sites, sometimes substantial, should not lead us to overestimate the trackmakers populations. These last were probably relatively moderately abondant in this inter-supratidal swamp environment. In the Causses, ichnites are connected with former algo-laminated deposits (Algal mats) which were rapidly hardened by means of calcitisation of cyanobacteria. The result has been a moderate depth of footprints; autopodia disturbing only a few cm of the carbonate substrate. Other fossils have been discovered : invertebrates with thin bivalve and gastropod shells, crustaceans tests and plants. These latter suggest the existence of paleomangroves like environments but also continental vegetation periodically overruning the swamp environment during regression/transgression cycles. At these times, wooded parts of it, could become protecting, feeding, resting and nesting places.

  

Avec ce volume se clôture l'étude géologique et paléontologique des sites à hominidés de Koobi Fora. Il fait suite aux deux précédents ouvrages, parus respectivement en 1978 et 1983, consacrés également à l'étude des faunes recueillies sur ces gisements dans le cadre du Koobi Fora Research Project. 

The subfamilies Murinae and "Dendromurinae" both include 4 species at Chorora. Among the former, while Preacomys nov. gen. seems to be a forerunner of Acomys, the affinities of the remaining, poorly known taxa, are more difficult to evaluate. The bulk of the fauna, remarkably, consists of Dendromurines. Their similarities with those of Ngorora tends to pull the site back in time, but the large size and diversity of Murines fit better an age more recent than the very beginning of the Late Miocene.

  

New records of terrestrial mammals from the Qasr el Sagha Formation, Fayum Depression, Egypt are reported, and the stratigraphic occurrences of these fossils noted. These include additional specimens of Moeritheríum, Barytherium, and anthracotheres, as well as the oldest record of a hyracoid in the Fayum.These Eocene mammals occur almost exclusively in the alluvial deposits of the Dir Abu Lifa Member of the Qasr el Sagha Formation and show close affinities to the faunas from the lower sequence of the Jebel Qatrani Formation. There is no evidence of a more marked faunal discontinuity between the Qasr el Sagha and Jebel Qatrani Formations than there is across any of the three major breaks in sedimentation that exist within the Jebel Qatrani Formation. The faunal similarities between fossils of the lower sequence of the Jebel Qatrani Formation and of the upper part of the Qasr el Sagha Formation is consistent with recent paleomagnetic dating that suggests that these rocks differ in age by only one to two million years. 

Extensive sampling of the Siwalik deposits of the Potwar Plateau of northem Pakistan and from the Zinda Pir dome of central Pakistan has produced a fossil record of Miocene ctenodactylids that can be correlated with the paleomagnetic time scale. The early Miocene Prosayimys flynni (n. gen., n. sp.) is recognized as the first ctenodactylid in the Indian subcontinent. Prosayimys is ancestral to Sayimys. From the late early Miocene to the early late Miocene, there is an anagenetic succession of three species of Sayimys: S. cf. S. intermedius, S. sivalensis, and S. chinjiensis (n. sp.). Sayimys chinjiensis gave rise to the late late Miocene S. perplexus. A second lineage is represented by Sayimys minor, S. sp. A, and S. sp. B. 

A fossil species of the extant genus Leuconoe, L. lavocati n. sp. from Le Garouillas Oligocene locality, Quercy phosphorites, SW-France, is established in nomenclatural standards. 

The review of material recently collected in the new localities from the “Phosphorites du Quercy", and different localities from the South of France, bring new informations on the genus Eucricetodon THALER. 1966, and Pseudocricetodon  THALER,  I969 (Middle and Upper Oligocene. Western Europe). Thanks to Eucricetodon huerzeleri VIANEY-LIAUD, 1972, which were unsufficiently known until now, is proposed. During the middle Oligocene Eucricetodon atavus  MISONNE, 1957 seems to give rise to two lineages. One of them led to Eucricetodon huberi,which however exhibits a larger size and a development of progressive characters on the teeth. The other would be Eucricetodon huerzeleri well differentiated at the “Mas de Pauffié" standard level (beginning of the upper Oligocene). The ornementation of lower incisors is described, when possible. Though the fossils are not abundant, it seems that the ancestral lineage, Eucriretodon atavus, remains in  the upper Oligocene (Boningen standard level). evolving into Eucricetodon praecursor SCHAUB, 1925 (Rickenbach standard level). The characters of Eucricetodon dubius  (SCHAUB. 1925), represented by a numerous population in Pech Desse and Pech du Fraysse (Quercy). confirm that this species and Eucricetodon praecursor  belong to two different lineages. As Eucricetodon dubius shows more primitive features, this species could not originale from Eucricetodon atavus -Eucricetodon huberi. The appearance of this species at the level of Mas de Pauffié could be the result of an immigration. A new definition of Pseudocricetodon incertus (SCHLOSSER. 1884) is given. This species has been found in several localities, where it had not been identified until now. lts comparison with Pseudocricetodon moguntiacus  (BAHLO. l975), found at several localities from the standard level of Antoingt (end of middle Oligocene). shows a parallel evolution to that of Pseudocetodon incertus, which is  of larger size and with a less complicated pattern of teeth. 

The revision of the old collections of fossil birds from the “Phosphorites du Quercy” and the study of new material give the following results (Gruiformes, Cariamae) :  The humeri and most of the carpometacarpi described under the name Filholornis belong in Elaphrocnemus. The ulnae ascribed to Fïlholornis belong in Idiornis. Most of the post-cranial elements of the genera Elaphrocnemus and Idiornis are described and show great similarities with recent Cariamidae and Opisthocomidae, and fossil Bathornithinae.  A new genus and a new species, Oblitavis insolitus, are created in the sub-family Idiornithinae; two new species are described in the genera Elaphrocnemus (E. brodkorbz) and Idiornis (I. itardiensis), and the species Elaphrocnemus gracilis is transferred to the genus Idiornis.  The genus Propelargus Lydekker is transferred from the family Ciconiidae to Cariamidae.  A new generic name, Occitaniavis, is created for the species Geranopsis elatus, which belong in Cariamidae, while the type-species of the genus, Geranopsis hastingsiae, is a member of the Gruidae.  The affinities between the Quercy avifauna and the Neotropical one is emphasized by the occurrence of Phorusrhacidae, previously known only from the Cenozoic of South America and the Late Pliocene or Early Pleistocene of North America. Thanks to the material collected during the new excavations, the stratigraphical position of most of the species is stated precisely, and evolutionary lineages are outlined. This study shows that the suborder Cariamae, presently restricted to two South American genera, was already extremely diversified during the Eocene, and widespread in Europe and North America. 

Nyctereutes megamastoides (Pomel) from the Villafranchian of the Auvergne and from Saint-Vallier presents cranial and dental characters sufficiently close to those of the late Pliocene canid from Perpignan (Roussillon), described by Depéret under the specific name of Canis donnezani belonging to the same genus Nyctereutes. The extinction of the European "Nyctereutes" group seems due to the too great alimentary specialization of this canid, whereas the Asiatic lineage represented in the Villafranchian by Nyctereutes sinensis Schlosser and at present by Nyctereutes procyonider Gray was able to maintain itself probably by means of a profound change in its alimentary regime.