The first good skeletal remains of Homogalax protapirinus from the Wasatchian of the Bighorn Basin, Wyoming, indicate that this primitive tapiromorph was more plesiomorphic in many features than primitive equoids including Hyracotherium. Compared to Hyracotherium, Homogalax more closely resembles Phenacodonta (the closest outgroup of Perissodactyla for which postcrania are known) in various details of articular surfaces, muscle attachments, and proportions of the humerus, manus, and pes.Among known taxa, Homogalax most nearly approximates the plesiomorphic postcranial skeletal anatomy of Perissodactyla. 

There is only one suid known in the Upper Miocene of Montredon (Hérault): Microstonyx (Limnostonyx nov. subgen.) antiquus (KAUP). It is differenciated from Microstonyx major by the presence of upper and lower canines which are considerably longer and biger. Its presence at Montredon corroborates the palustrine habitat for the species. 

 Detailed anatomical description of arsinoithere cranial remains from the Lower Oligocene, Fayum Depression, Egypt, provides the basic data for a systematic investigation. All cranial and some postcranial features are assessed from a phylogenetic standpoint. Several soft tissue characters are then added to a cladistic analysis based on 54 derived ungulate morphological characters. The resulting phylogenetic hypothesis implies that perissodactyls, sirenians, proboscideans and arsinoitheres constitute a monophyletic unit (5 synapomorphies). However, increasing the tree length by 3 steps reveals a closer association between hyraxes and perissodactyls. Nevertheless, 13 synapomorphies link proboscideans, sirenians and  arsinoitheres to the exclusion of all other ungulates. Form of the sphenopalatine and ethmoid foramina, recurved posttympanic process, absence of a fenestra rotundum in the petrosal, vestigial paroccipital process of the exoccipital and the highly unusual absence of a hypoglossal foramen in the skull, imply a robust sister-group relationship between arsinoitheres and proboscideans. In this analysis artiodactyls share only one derived character with all other ungulates studied. Monophyly of Ungulata, including Artiodactyla, is therefore only weakly supported. It is argued that pedal anatomy of hyraxes is non-homologous with that of Tethytheria. Arsinoitherium should now be classified within Tethytheria, sharing a sister-group relationship with Proboscidea. Hyraxes are excluded, thus refuting the concept of Paenungulata. However, monophyly of the wider concept, Pantomesaxonia, containing hyraxes, perissodactyls, sirenians, proboscideans and now, arsinoitheres, is supported by this study. 

In this article is described a new species of Nyctitheriidae with primitive characters: Leptacodon nascimentoi n. sp. from the early Eocene of Silveirinha (Portugal). 

Abstract: A very tiny cuckoo, Chambicuculus pusillus, was recently described after a few fragments of tarsometatarsi from the late Early/early Middle Eocene of Djebel Chambi, Tunisia. New remains, notably a coracoid, confirm the attribution of this genus to the recent family Cuculidae. This coracoid shows a very elongate and strap-like processus procoracoideus. This morphological feature is otherwise only known in the family Cuculidae. The characteristics of the coracoid and tarsometatarsi show that Chambicuculus is morphologically more advanced over the other stem cuculids described in Europe and North America. Chambicuculus is the oldest Cuculidae known so far. 

An ilium from the Upper Eocene (MP l7a) of Hordle, England, represents the first report of Rana from the Eocene of Britain. The ilium is similar to those of the water frog (Rana [ridibunda]) species group. 

The Squamates from Sainte-Néboule are charateristic of the Upper Eocene from the Phosphorites du Quercy. Most genera known from this locality do not survive the « Grande Coupure ». Cadurcoboa insolita gen. nov., sp. nov. is a small Boidae very characteristic of the Upper Eocene. 

The principles and practices employed in establishment and recognition of South American land mammal ages are reviewed along with previous and present concepts of distinguishing time, rock, and faunal units. Previous chronological arrangements of South American Tertiary land mammal faunas are appraised on the basis of recent geological and paleontological data. Twelve South American Tertiary land mammal ages are here recognized [from oldest to youngest, Riochican (middle to late Paleocene); Casamayoran (early Eocene); Mustersan (middle Eocene); Divisaderan (late Eocene); Deseadan (early [to middle?] Oligocene); Colhuehuapian (late Oligocene); Santacrucian (early Miocene); Friasan (middle Miocene); Chasicoan (late Miocene); Huayquerian (latest Miocene); Montehermosan (early to middle Pliocene); and Chapadmalalan (late Pliocene)]. As all except the Friasian were originally defined on the basis of Argentine faunas, these are discussed first and at length, and each is reviewed with discussion of type locality, stratigraphy, type fauna, and faunal correlations. Non-Argentine faunas are then discussed country by country in alphabetical order.

     A review is given of radioisotope dates obtained on volcanic rocks (i.e., basalts, tuffs) associated with mammalbearing beds in Argentina. Based on these age determinations and on correlation of the late Tertiary land mammals involved in the interchange between North and South America, a chronology of South American land mammal ages correlated with North American land mammal ages and European marine stages is proposed.

It is concluded that South America was an island continent through most of the Tertiary Period (ca 65 to about 3 Ma). As a result, the land mammal fauna of South America developed in isolation and was dominated by autochthonous endemic groups. Toward the end of the Tertiary (i.e., middle Miocene) a unique faunal balance had been achieved by the descendants of the ancient inhabitants (notoungulates, litopterns, condylarths, astrapotheres, edentates, marsupials) and of later (late Eocene) waif immigrants (caviomorph rodents, platyrrhine primates). A prominent feature of this mammal fauna was the combination of carnivorous and omnivorous marsupials with native placental herbivorous ungulates, subungulates, and edentates.

Sometime during the late Miocene, a limited but important interchange of mammalian taxa between North and South America took place. Procyonids (raccoons and their allies), a group of North American origin, first appear in South America in strata of Huayquerian Age, while members of the extinct South American ground sloth families Megalonychidae and Mylodontidae first appear in North America in early Hemphillian time. These groups dispersed along island arcs before the appearance of the Panamanian land bridge in the Pliocene (ca 3.0 Ma). Cricetine rodents, a group of North American origin, are first known in South America in strata of Montehermosan Age. The known taxa are too advanced and diversified to be considered the first of this group to invade South America. lt is believed by some workers that these rodents arrived before the Montehermosan, possibly in the late Miocene or earlier, by waif dispersal from North America.

The isolation of South America ended with the appearance of the Panamanian land bridge, which provided a direct, dry land connection between the two Americas. Across this portal an extensive interchange of terrestrial faunas occurred, and the fossil record documents an intermingling of these long-separated land mammals faunas.

      The beginning of this interchange by land route in South America is marked by the appearance of mammals which evolved from North American emigrants in the Chapadmalal Formation of Argentina. These include a mustelid (Conepatus), a tayassuid (Argyrohyus), and four genera (Akodon, Dankomys, Graomys, Reithrodon) of cricetine rodents. The appearance of this contingent of northern animals favors the existence of the Panamanian land bridge by this time. Likewise, a large number of terrestrial vertebrates of South American origin appear in North America in beds of late Blancan Age date around 2.7 Ma. Among the mammals are Neochoerus, Erethizon, Glyptotherium, Glossotherium, Kraglievichia, and Dasypus. 

Fossils of the Trionychidae, Bataguridae or Emydidae, Palaeophis and Crocodylia from Chattahoochee, NW Florida, USA, represent the first report of an Eocene reptile fauna from Florida. 

Deux types de remaniements post mortem me paraissent caractéristiques de l'état de conservation des dents de mammifères fossiles dans les Phosphorites du Quercy :

1) Des destructions localisées d'origine biologique, sous forme de galeries de morphologie très variable creusées dans la dentine et le cément, et impliquant sans doute la participation de différents types de micro-organismes. Ces altérations se sont développées peu de temps après la mort, avant la fossilísation proprement dite et se sont rapidement arrêtées après l'enfouissement dans le sédiment phosphaté.

2) Des perturbations dans les structures de la dentine liées aux variations locales de minéralisation, provoquées par une imprégnation diffuse des zones les moins calcifiées par divers minéraux et probablement surtout de l`apatite. 

Record of life and works of Henri Menu, French zoologist, contributor to the knowledge of living and fossil bats. 

Aside from Madtsoiidae, anilioids, and Boidae that were studied previously, the middle Palaeocene of ltaborai (BraziI) has produced Ungaliophiinae ("tropidophiids"), booid-grade snakes incertae sedis, and a possible Russellophiidae (Caenophidia) that are described in the present article. This article is the third and final report on the snakes from the locality. The Ungaliophiinae (Paraungaliophis pricei gen. et sp. nov.) are rare whereas the booid-grade snakes incertae sedis (ltaboraiophis depressus gen. et sp. nov., Paulacoutophis perplexus gen. et sp. nov.) are more frequent. A single vertebra is referred to the Russellophiidae (Caenophidia) with reservation. An update of the whole fauna of snakes from ltaborai is provided. Hechtophis austrinus that was tentatively referred to the erycine Boidae is now regarded as a Boidae incertae sedis. Most snakes from Itaborai are known only from the locality. Astonishingly, only the ailioids Coniophis cf. C. precedens gives possible evidence of interchanges between South and North America. The fauna of snakes from Itaborai, as well as the other Palaeocene faunas of snakes from South America are distinct from those of the Cretaceous and the Eocene of South America; they appear to be more different from the Cretaceous faunas than from those of the Eocene. The fauna from Itaborai is the richest and most diverse assemblage of snakes from the Palaeocene worldwide; it shares only a few taxa with other Palaeocene localities. 

This paper is mainly devoted to the systematics of rodents from two middle Eocene (Bartonian) localities: Chéry-Chartreuve and Rocourt-Saint-Martin (Aisne, Eastern Paris Basin). These two localities are stratigraphically located slightly above the Auversian sands. The two faunas, which comprise 11 and 8 taxa, respectively, are very different in their composition. That of Rocourt-Saint-Martin shows strong similarities with that of the geographically very close locality of Grisolles, referred to the MP16 mammalian Reference level. The very distinct fauna of Chéry-Chartreuve includes a new species of Ailuravinae, Ailuravus nov.sp, and some teeth of the theridomyid Protadelomys, which represent archaic elements in the fauna. The most abundant species of the locality represents a new genus of primitive Theridomyidae. The presence of some teeth belonging to a new species of large Remyinae, Remys nov. sp., of Elfomys engesseri HOOKER & WEIDMANN, and a population of small dimensions referred to the genus Estellomys allow a correlation with Les Alleveys (Switzerland), with however some differences that would indicate an older age for Chéry-Chartreuve. Situated at the base of the "Marinesian" from the Bassin de Paris, this fauna is unquestionably different from those referred to the MP16 reference level and could represent an older level for which the macrofauna remains very poorly known. Conversely, the comparison of rodents from La Livinière II with those present in MP16 faunas, especially those of Robiac (Gard), shows a great similarity between both localities. This casts doubts on whether to keep this La Livinière II faunule to define the current MP15 reference level, while the biostratigraphical position of Pontils 26 (Spain), previously referred to this level, is reconsidered. Chery Chartreuse could be a good candidate for a new definition of the MP15 reference level. 

Prolagus sardus is the last representative of the diverse lineages of European endemic ochotonids. It is also the most abundant in the collections. The previous studies made of this species have established rather well its dental morphology, its phylogenetic position, its geographic and temporal distribution, and its intraspecific individual variation. On the other hand, no osteologic study has fully utilized the superb material from Corsica and Sardinia  collected by Forsyth Major.
Nearly all of the parts of the skeleton are represented in this material; they are here described and figured. Comparisons are made with Ochotona, the only surviving genus of the family, as well as with the living leporids, in particular Romerolagus, Oryctolagus and Lepus. A brief examination of the dentition reveals some particular characters of the incisors. The osteologic study allows limited interpretations to be advanced concerning the posture, mastication. locomotion and some other adaptive features of P. sardus.
P. sardus appears as an incontestable ochotonid, but it differs from Ochotona by some characters which are found in leporids. Certain of the characters are here judged primitive for lagomorphs. The genus Ochotona therefore can no longer be considered as the image of a primitive lagomorph, because in spite of the lack of cursorial adaptations which differentiate it from leporids, it presents other specializations which are common to it alone. 

This paper is about the oldest African rodents faunas, from the late Early Eocene, or early Middle Eocene, Glib Zegdou (Algeria) and Chambi (Tunisia) localities. Five species are described and figured, belonging to a new family here created, the Zegdoumyidae.

This family is compared to the Early and Middle Eocene rodents families from Asia, Europe and North America (Chapattimyidae, Yuomyidae, Gliridae, Theridomyidae, lschyromyidae and Sciuravidae), as well as to those known from the Late Eocene African locality Bir El Ater (Anomaluridae and Phiomyidae).

On the one hand, it seems clear that the African endemic Anomaluridae arise from the Zegdoumyidae. On the other hand, the lschyromyidae, or primitive Sciuravidae, may be the most reliable ancestral groups for the Zegdoumyidae. Thus, this new family can be considered as the sister group for the American Sciuravidae on the one hand, and for the European Gliridae on the other hand.

The biogeographical consequences of these phylogenetic hypotheses are discussed. A new phase of communication between Europe and North Africa is inferred, during the Early Eocene. It has been followed by a short period of endemism, allowing the adaptive radiation for the Zegdoumyidae, preceding the immigration of the Phiomyidae, during the Late Eocene, probably from Asian relatives. 

Abundant crocodilian fossil remains have been collected, mainly by R. Lavocat. from several levels of the
Palaeogene of the Tilemsi valley (eastern Mali). Several dyrosaurids occur in the Palaeocene (Montian ?) : Phosphatosaurus sp., Hyposaurus bequaerti (Dollo). H. wilsoni (Swinton). H. nopesai (Swinton), Rhabdognathus rarus Swinton, R. compressus n. sp. ln the lower Eocene has been found a dyrosaurid which cannot be accurutely determined.In the middle Eocene were found a dyrosaurid, Tilemsisuchus lavocati n. g., n. sp., and an indeterminate eusuchian. 

The dentition of Welcommia bodeuri nov. gen. nov. sp. from the Valanginian of Southem France is described and reconstructed. Species and genera of Upper Jurassic and Lower Cretaceous Hexanchiformes are reviewed and discussed.
The genus Notidanoides MAISEY, 1986 must be restricted to the single Nusplingen Upper Jurassic specimen, whose attribution to the species muensteri AGASSIZ, 1843 remains doubtful.
The genus Paranotidanus WARD &THIES, 1987 that does not rest on any type-species nor on any precise dental characterization must be rejected.
The genus Eonotidanus PFEIL, 1983, based on a very poorly preserved and heterogeneous type-material must also be rejected.
Teeth from the Lower Cretaceous, with a peculiar morphology, previously assigned to Eonotidanus or to Notidanoides, are to be ranked in the new genus Pachyhexanchus. 

Comparison of various measurements of taxonomic evolution using stratigraphic range data for orders, families and genera of land mammals indicates several means by which deficiencies of the South American fossil record (e.g., presence of hiatuses, unequal temporal and geographic representation of ages, unequal systematic treatment) may be normalized, thus permitting a less distorted appreciation of diversity pattern and trend. Initial radiation of native taxa resulted in a relative equilibrium by early Eocene time. Subsequent increases in absolute diversity were apparently induced by immigration at the family level and by environmental factors at the generic level. Miocene through Pleistocene phases of faunal stability, herein characterized as chronofaunas, are punctuated by rapid turnover events resulting from a complex of factors, including adaptive radiation of immigrant taxa into unoccupied eco-space; environmental and concomitant habitat change induced by orogenic events of the Andes; and biotic interactions between native and immigrant taxa, including competition and prey naivete. The first two factors account for major faunal transitions in the South American middle and late Tertiary; immigration-induced turnover may have been of greater importance in shaping the character of the fauna upon the Great American Interchange and the arrival of man in the Neotropics 

Pleistocene bird fossils have been studied from nine localities on Crete. Part of this material was described earlier by the author (Weesie, 1982) and will not be treated here in extenso, the results will be incorporated. More than one third of the over 10,000 fossil bird bones available could be identified ; they were found to represent at least 65 bird species. The following species of the Pleistocene Cretan avifauna are new to the fauna of Crete : Branta ruficollis, Haliaeetus albicilla, Gyps melitensis, Aquila chrysaetos simurgh n. ssp., Ketupa zeylomensis, Aegolius funereus, Dendrocopos leucotos, Zoothera dauma, Turdus iliacus and Pyrrhula pyrrhula. The Pleistocene Cretan avifauna differs less from comparable mainland avifaunas than (fossil) avifaunas from oceanic islands do. Still, the Pleistocene Cretan avifauna has two qualities that are characteristic of island avifaunas : the almost complete absence of a group of birds (the Galliformes) and the presence of two endemic (sub)species : the giant eagle Aquila chrysaetos simurgh n. ssp. and the long-legged owl Athene cretensis (Weesie, 1982). The new subspecies is described in the present study.
These endemic birds of prey were found in association with their supposedly principal prey species (now extinct as well) : endemic mice for the owl and endemic deer for the eagle. Endemic mammals have been found in association with endemic birds of prey on many islands, not only in the Mediterranean. There is evidence that the size of endemic birds of prey becomes optimally adapted to their feeding on certain endemic mammals, especially rodents. Another characteristic of the Pleistocene Cretan avifauna is the great number of species of birds of prey. This appears to be a common characteristic of fossil avifaunas from caves on Mediterranean islands as well as from caves on the European mainland. However, we think that ecological conditions on Pleistocene Crete (especially the abundant presence of mice) helped to account for the high representation of birds of prey. Furthemore, the fossil avifauna enables us to draw some conclusions about the climate and vegetation on Pleistocene Crete : it is concluded that the climate was cooler than today and that Crete was largely covered with forests. Finally, the reasons for the extinction or disappearance from Crete of some bird species of the Pleistocene Cretan avifauna are discussed. 

This paper is an English concentrate of various Russian publications by the senior author presenting the mammaIian taxa from the Cretaceous (Albian through Santonian) of the region termed Middle Asia by Soviet geographers. The diagnoses are the unmodified, literal translation of the original version, but are followed with short complementary remarks; most of the species are illuslrated anew with SEM photographs, others are by normal photography. The fossiliferous formations are cited and arguments for their dating are given. Finally, the main vertebrate groups accompanying mammaIs are listed and the environment and climate at the time of deposition are suggested. In conclusion, an hypothesis on the origin and high diversity of tribosphenic mammals on the Cretaceous coastal plains of southwest Asia is proposed. In appendix the taxon Khuduklestes bohlini novo gen. novo sp. is formally defined.